Personally I think it is even absurd that someone needs to point this out in a video! As a kid in South America I used to walk 80% of the time barefoot or on sandals. It is clear that in such situation any faster movement needs to be done toe first. So, fighting, dancing, running, would be done toe first in such cultures. On the other hand, heel-first is more energy efficient, so you do that when walking slowly barefoot. It was modern shoes that allowed people to do everything heel-first.
We moved back to NZ where I grew up, and our kids went to a 50-person primary school with not a pair of shoes in sight. You learn where the patches of prickles are and avoid those bits of the grass. (Data point: the prickles are in the same places at the beach and community hall as they were 40 years ago when I were a lad)
The purpose of the shoot was to determine whether a galloping horse ever lifts all four feet completely off the ground during the gait; at this speed, the human eye cannot break down the action.
That’s ridiculously interesting, but is it really comparable to a walking speed human? Sounds more like an issue with the human eye than older paintings in general
If you look at the middle right image and compare it to the painting it's not all that wrong. Note: horse on right it jumping, not simply running. @ 20 seconds: https://youtu.be/oLvAxzVeyLs