then why 5400k? At work (90% mac shop) we are just looking at what to do for a central server. Apple isn't supporting that market very well - they need a shuttle sff size mac midi that uses full fat non mobile parts.
Honestly, it could be due to heat issues. It's a small device and faster drives can get hot.
Apple has tried to support that market before, and it's just not a big market, and it isn't a one-size fits all market. Many people have been clamoring for an Apple solution for SMB server, and it just isn't profitable. This is especially true when you take into account cloud shared storage providers like Dropbox or Box.net.
It's also a market that is more focused on margins. It's easier to charge a higher margin for a device you use everyday, but what about one that you install in a closet or put in a corner and forget?
An HP microserver running FreeNAS is a tough combination to beat for an SMB central server.
It can't be due to heat issues, because Apple sells this exact same machine in "server" spec, with 2 HDDs stacked on top of each other, so if that enclosure can deal with 2xHDDs then it certainly can deal with 1x7200rpm hdd.
It looks like it's a similar design externally, but it might have changed internally. For example, there are two thunderbolt ports now. They also don't sell the "server" configuration anymore, which had also had 5400rpm drives.
So, it still could be from heat. It's probably a cost issue, but there is a lot at play in such a small system.